Staff Fellow, Lab Director: Robert London, National Institutes of Health, National Institute of Environmental Health Science, Research Triangle Park, NC, 1989-1990.

Senior Staff Fellow, Lab Director: Robert London, National Institutes of Health, National Institute of Environmental Health Science, Research Triangle Park, NC, 1990-1992.

Research Assistant Professor, Ophthalmology/Radiology/Biomedical Engineering, University of Texas Southwestern Medical Center, Dallas, TX, 1992-1996.

Associate Professor, Anatomy/Cell Biology and Kresge Eye Institute, Wayne State University School of Medicine, Detroit, MI, 1996-2002.

Professor, Anatomy/Cell Biology and Kresge Eye Institute, Wayne State University School of Medicine, Detroit, MI, 2002-present.